Index fossil for the Silurian Period Waldron Shale thought to be 425 to 411 million years old. These crinoids must have flourished on the seafloor. Judging by their root structure, the seafloor was pretty soft allowing the animal to create extensive holdfasts. See my friend Dave's July 24, 2011 posting at Views of the Mahantango on holdfasts CLICK HERE.
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